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Crudup starred in the Netflix original series Gypsy in 2017. In 2019, he began playing Cory Ellison in the Apple TV+ drama series The Morning Show, which earned him the Critics' Choice Television Award for Best Supporting Actor in a Drama Series and a nomination for the SAG Award for Outstanding Performance by a Male Actor in a Drama Series.

In November 2017, he starred in the world premiere of David Cale's one-man play Harry Clarke at Vineyard Theatre. It moved to the Minetta Lane Theatre the following spring.

Crudup has been in a relationship with actress Naomi Watts since 2017.

In August 2013, he co-starred with Ian McKellen and Patrick Stewart in the Harold Pinter play No Man's Land as well as in Waiting for Godot at the Berkeley Repertory Theatre. The shows transferred to The Cort Theatre in New York City, where they ran in repertory until March 2014.

In 2011, Crudup received a Tony Award nomination for Featured Actor in a Play for his role in the Broadway revival of Arcadia.

He starred in The Metal Children, an off-Broadway play written and directed by Adam Rapp in 2010.

He appeared as Zartan in the 2009 parody video The Ballad of G.I. Joe on the website Funny or Die.

Crudup completed filming Watchmen with director Zack Snyder in Vancouver, British Columbia. He portrayed the superhero Doctor Manhattan. He portrayed former U.S. Secretary of the Treasury Timothy Geithner in a TV film about 2008's financial crisis, Too Big to Fail (2011).

Crudup received a 2002 Tony Award nomination for Best Actor in a Play for his performance as the title character in The Elephant Man on Broadway, as well as a 2005 nomination for his role as Katurian in the Broadway production of The Pillowman, also starring Jeff Goldblum, which closed on September 18, 2005. From October 2006 through May 2007, he was featured in the first two parts of The Coast of Utopia by Tom Stoppard at Lincoln Center, playing literary critic Vissarion Belinsky, for which he received a 2007 Tony Award for Best Featured Actor in a Play.

From 1998 to 2005, Crudup was the narrator for the U.S. television ad campaign "Priceless" for MasterCard. In the ads, the narrator (Crudup) lists the prices of two goods or services, then lists some third, intangible benefit gained from those purchases and concludes, "priceless". He said in 2005 that appearing in the ads "changed my life", in that they gave him the financial freedom to pursue the acting work that he wanted to do.

Crudup began acting in films such as 1996's Sleepers, 1997's Inventing the Abbotts, and 1998's Without Limits, where he played the role of running legend and Olympian Steve Prefontaine. His first role in an animated feature was in 1999's English release of Princess Mononoke, in which he starred as Ashitaka. He then played Russell Hammond, the lead guitar player of the fictional band Stillwater in Cameron Crowe's Almost Famous (2000). In 2006's The Good Shepherd, he played British spy Arch Cummings, a stand-in for Kim Philby. The same year, he played a supporting role in Mission: Impossible III. In 2007, he played the leading role of Henry Roth in the film Dedication.

From 1996 to November 2003, Crudup dated actress Mary-Louise Parker. Parker was seven months pregnant when Crudup ended their relationship and began dating actress Claire Danes. Crudup and Danes ended their relationship in 2006.

Crudup attended the University of North Carolina at Chapel Hill, where he received an undergraduate degree, and he continued his passion for acting with the undergraduate acting company, LAB! Theatre. He also acted for UNC-STV's most popular show, General College. He was a member of the Beta Chapter of Delta Kappa Epsilon. He then studied at New York University's Tisch School of the Arts graduate acting program, where he earned a Master of Fine Arts degree in 1994.

Crudup was born in Manhasset, New York. His parents, Georgann (née Gaither) and Thomas Henry Crudup III, divorced during his childhood, and later remarried, before divorcing a second time. On his father's side, he is a descendant of Congressman Josiah Crudup of North Carolina. His maternal grandfather was William Cotter "Billy" Gaither, Jr., a well-known Florida trial lawyer, and his maternal grandmother later remarried to Episcopal bishop James Duncan. Born second of three boys, Crudup has two brothers: Tommy, an executive producer, and Brooks, also a producer. He left New York with his family when he was about eight years old, first living in Texas, then in Florida. He graduated from Saint Thomas Aquinas High School in Fort Lauderdale, Florida in 1986.

William Gaither Crudup (/ˈ k r uː d ə p / born July 8, 1968) known professionally as Billy Crudup, is an American actor. He is a four-time Tony Award nominee, winning once for his performance in Tom Stoppard's The Coast of Utopia in 2007. He has starred in numerous high-profile films, including Without Limits, Princess Mononoke, Almost Famous, Big Fish, Mission: Impossible III, Watchmen, Public Enemies, Spotlight, Jackie, The Stanford Prison Experiment, Justice League, and Alien: Covenant, in both lead and supporting roles. He has been nominated for an Independent Spirit Award for Best Male Lead for his performance in Jesus' Son, and received two Screen Actors Guild Award nominations as part of ensemble cast for Almost Famous and Spotlight, winning for the latter.