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Fletcher's debut as a director was for a script he co-wrote, Wild Bill, was released on 20 March 2012. His second film as director is a musical film by Stephen Greenhorn, Sunshine on Leith based around the popular Proclaimers songs which was released on 4 October 2013. In 2015, he directed the feature film Eddie the Eagle. On 6 December 2017, Fletcher was announced as Bryan Singer's replacement director on the Queen biopic, Bohemian Rhapsody. The film was released on 2 November 2018. While Fletcher had helped finish the film, Singer received sole directing credit. Fletcher's latest directing role was on Rocketman, a biopic about the life of Elton John. His next project is going to be Sherlock Holmes 3, the sequel to the 2011 film Sherlock Holmes: A Game of Shadows, expected to release December 21, 2021.

Fletcher made his directorial debut with Wild Bill (2011), and also directed Sunshine on Leith (2013) and Eddie the Eagle (2015). He replaced Bryan Singer as director of Bohemian Rhapsody, a biopic about the band Queen, released in October 2018; due to DGA rules, he received executive producer credit. In 2019 he directed Rocketman, a film based on the life of Elton John.

2011 saw him acting in his first role as a police officer, as Detective Edwards in the British film noir Jack Falls, alongside his Lock, Stock and Two Smoking Barrels co-stars Jason Flemyng and Alan Ford. In 2012, Fletcher had a small role in horror-comedy Cockneys vs Zombies, where he played the character 'Dad Maguire'. In 2015 Fletcher played the role of 'Scooter' in episode 5 of the 8-part BBC TV drama The Interceptor. He also took the lead role as history teacher Mr Sheers in supernatural British teen film Coven. In 2016 Fletcher played the role of 'Paul' in the cult British crime comedy, Smoking Guns.

He also appeared in the Boon episode "Walking Off Air", playing Eddie Cotton. In 2009, he appeared in Misfits as Nathan Young's dad, reprising the role in 2010 for the second series. Fletcher stars in 2009 in the vampire film Dead Cert, which is directed by Steven Lawson.

On television Fletcher has appeared in the major HBO drama, Band of Brothers and in a supporting role in the BBC One historical drama The Virgin Queen (US PBS 2005, UK 2006). He also appeared in Kylie Minogue's music video for "Some Kind of Bliss" (1997). He starred on BBC One in a series based on Imogen Edwards-Jones's book Hotel Babylon that ran for four series before being cancelled in 2009. He also appeared in "The Booby and the Beast", an episode in the second series of the BBC's series Robin Hood and in the 2008 radio series The Way We Live Right Now. He appeared in the Bo' Selecta! spinoff A Bear's Tail as The Scriptwriter. He played a brief role in the BBC series New Tricks, in the episode "Final Curtain", as an actor named Tommy Jackson.

In 1998, Fletcher featured on the song "Here Comes the Flood" from the album Fin de Siecle by The Divine Comedy. In 2014 he narrated the BBC1 show Del Boys and Dealers.

Fletcher was born in Enfield, England to parents who were teachers and grew up in Woodford Green and Palmers Green. He dated his Press Gang co-star Julia Sawalha and also had a relationship with actress Liza Walker. In 1997 he married Lithuanian film and theatre director Dalia Ibelhauptaitė in Westminster. His best man was fellow actor Alan Rickman. Dexter's brothers are also actors; Graham Fletcher-Cook and Steve Fletcher.

Fletcher recalls presenting GamesMaster was 'very unforgiving' and 'quite intimidating'. He claimed the most difficult part was 'just getting up, taking a deep breath, and going out there and doing it'. His style of presenting was heavily criticised by viewers who said his 'in your face' method was too 'OTT' and did not suit the show. Some viewers suspected his genuine Cockney accent was put on for effect. The series finished in 1994 and Dominik Diamond returned for the last four series until the show ended in February 1998. The show's creator Jane Hewland later said in a retrospective interview, that she sees the third series as a "lost" one, due to admitting to a series of poor decisions and mistakes, including hiring Fletcher as main presenter. His name was however included on a rolling list of acknowledgements, which replaced the usual end credits when the final episode was broadcast, but he has not taken on any other presenting role before or since.

Fletcher presented the third series of Channel 4 show GamesMaster in 1993. Dexter was originally slated to be a special guest on the third series but was then asked to be presenter, after Dominik Diamond left the show due to his disapproval at McDonald's landing a sponsorship deal with them.

Fletcher has been the voice for McDonald's television adverts and (feigning a US accent) is the narrator of The Game audio book written by Neil Strauss. He also narrated the Five series Airforce Afghanistan, as well as the Chop Shop: London Garage series on the Discovery Channel. In 1993, he was the voice of Prince Cinders in the short animated comedy of the same name. Also in 1993, he was the uncredited UNIT soldier narrator of the UNIT Recruiting Film - a five-minute spoof piece that preceded a BBC1 repeat of the sixth and final episode of Doctor Who story Planet of the Daleks.

Fletcher trained at the Anna Scher Theatre. His first film part was as Baby Face in Bugsy Malone (1976). He made his stage début the following year in a production of A Midsummer Night's Dream. As a child actor he was regularly featured in British productions in the early 1980s, including The Long Good Friday, The Elephant Man and The Bounty. In 1987 Fletcher was cast in Lionheart. As an adult he appeared on television as the rebellious teenager Spike Thomson in Press Gang and in Murder Most Horrid (1991) with Dawn French. He has also starred in the films Caravaggio (1986), The Rachel Papers (1989), Lock, Stock and Two Smoking Barrels (1998), Layer Cake (2004), AffirmFilm's Solomon as Rehoboam and Universal's Doom.

Dexter Fletcher (born 31 January 1966) is an English actor and director. He has appeared in Guy Ritchie's Lock, Stock and Two Smoking Barrels and the crime comedy Smoking Guns, as well as in television shows such as the comedy-drama Hotel Babylon, the HBO series Band of Brothers and, earlier in his career, the children's show Press Gang. He was also in the film Bugsy Malone. He had a short stint as a presenter on the third series of Channel 4's GamesMaster in 1993 and 1994.