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On August 11, 2015, Netflix announced that Mascots, a film directed by Guest about the competition for the World Mascot Association championships's Gold Fluffy Award, would debut in 2016.

In 2013, Guest was the writer and producer of the HBO series Family Tree, a lighthearted story in the style he made famous in Spinal Tap, in which the main character, Tom Chadwick, inherits a box of curios from his great aunt, spurring interest in his ancestry.

In 2010, the United States Census Bureau paid $2.5 million to have a television commercial directed by Guest shown during television coverage of Super Bowl XLIV.

He is also currently a member of the musical group The Beyman Bros, which he formed with childhood friend David Nichtern and Spinal Tap's current keyboardist C. J. Vanston. Their debut album Memories of Summer as a Child was released on January 20, 2009.

Guest holds a hereditary British peerage as the 5th Baron Haden-Guest, and has publicly expressed a desire to see the House of Lords reformed as a democratically elected chamber. Though he was initially active in the Lords, his career there was cut short by the House of Lords Act 1999, which removed the right of most hereditary peers to a seat in the parliament. When using his title, he is normally styled as Lord Haden-Guest. Guest is married to actress and author Jamie Lee Curtis.

The experience of making Spinal Tap directly informed the second phase of his career. Starting in 1996, Guest began writing, directing and acting in his own series of substantially improvised films. Many of them came to be definitive examples of what came to be known as "mockumentaries"—not a term Guest appreciates in describing his unusual approach to exploring the passions that make the characters in his films so interesting. He maintains that his intention is not to mock anyone, but to explore insular, perhaps obscure communities through his method of filmmaking.

Guest became the 5th Baron Haden-Guest, of Great Saling, in the County of Essex, when his father died in 1996. He succeeded upon the ineligibility of his older half-brother, Anthony Haden-Guest, who was born prior to the marriage of his parents. According to an article in The Guardian, Guest attended the House of Lords regularly until the House of Lords Act 1999 barred most hereditary peers from their seats. In the article Guest remarked:

Guest appeared as Dr. Stone in A Few Good Men (1992), as Lord Cromer in Mrs Henderson Presents (2005) and in the 2009 comedy The Invention of Lying.

Upon his father succeeding to the family peerage in 1987, he was known as The Hon. Christopher Haden-Guest. This was his official style and name until he inherited the barony in 1996.

He appeared as Count Rugen in The Princess Bride. He had a cameo role as the first customer, a pedestrian, in the 1986 musical remake of The Little Shop of Horrors, that also featured Steve Martin. As a co-writer and director, Guest made the Hollywood satire The Big Picture.

Guest's biggest role of the first two decades of his career is likely that of Nigel Tufnel in the 1984 Rob Reiner film This Is Spinal Tap. Guest made his first appearance as Tufnel on the 1978 sketch comedy program The TV Show.

Along with Martin Short, Billy Crystal and Harry Shearer, Guest was hired as a one-year only cast member for the 1984–85 season on NBC's Saturday Night Live. Recurring characters on SNL played by Guest include Frankie, of Willie and Frankie (coworkers who recount in detail physically painful situations in which they have found themselves, remarking laconically "I hate when that happens"); Herb Minkman, a shady novelty toymaker with a brother named Al (played by Crystal); Rajeev Vindaloo, an eccentric foreign man in the same vein as Andy Kaufman's Latka character from Taxi; and Señor Cosa, a Spanish ventriloquist often seen on the recurring spoof of The Joe Franklin Show. He also experimented behind the camera with pre-filmed sketches, notably directing a documentary-style short starring Shearer and Short as synchronized swimmers. In another short film from SNL, Guest and Crystal appear as retired Negro league baseball players, "The Rooster and the King".

Guest married actress Jamie Lee Curtis in 1984 at the home of their mutual friend, Rob Reiner. They have two adopted children: Anne (born 1986) and Thomas (born 1996). Because Guest's children are adopted, they cannot inherit the family barony under the terms of the letters patent that created it, though a 2004 Royal Warrant addressing the style of a peer's adopted children states that they can use courtesy titles. The current heir presumptive to the barony is Guest's younger brother, actor Nicholas Guest.

Guest played a small role in the 1977 All In the Family episode "Mike and Gloria Meet", where in a flashback sequence Mike and Gloria recall their first blind date, set up by Michael's college buddy Jim (Guest), who dated Gloria's girlfriend Debbie (Priscilla Lopez).

Guest studied acting at New York University's Graduate Acting Program at the Tisch School of the Arts, graduating in 1971.

Guest began his career in theatre during the early 1970s with one of his earliest professional performances being the role of Norman in Michael Weller's Moonchildren for the play's American premiere at the Arena Stage in Washington D.C. in November 1971. Guest continued with the production when it moved to Broadway in 1972. The following year he began making contributions to The National Lampoon Radio Hour for a variety of National Lampoon audio recordings. He both performed comic characters (Flash Bazbo—Space Explorer, Mr. Rogers, music critic Roger de Swans, and sleazy record company rep Ron Fields) and wrote, arranged and performed numerous musical parodies (of Bob Dylan, James Taylor and others). He was featured alongside Chevy Chase and John Belushi in the Off-Broadway revue National Lampoon's Lemmings. Two of his earliest film roles were small parts as uniformed police officers in the 1972 film The Hot Rock and 1974's Death Wish.

Christopher Haden-Guest, 5th Baron Haden-Guest (born February 5, 1948) is an English-American screenwriter, composer, musician, director, actor, and comedian. Guest is most widely known in Hollywood for having written, directed and starred in his series of comedy films shot in mock-documentary (mockumentary) style. Many scenes and character backgrounds in Guest's films are written and directed, although actors have no rehearsal time and the ensemble improvise scenes while filming them. The series of films began with This Is Spinal Tap (which he did not direct) and continued with Waiting for Guffman, Best In Show, A Mighty Wind, For Your Consideration, and Mascots.