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In January 2008, Speight found Collins's body in the bath of their shared London flat. He was arrested on suspicion of her murder, but not charged with any offence. An inquest later determined that Collins had died of a drug overdose and severe burns from hot water. In April that year, Speight was reported missing and was later discovered to have hanged himself near Paddington station. Two suicide notes were discovered, describing how he could no longer live his life without Collins.

Speight was born in Seisdon, Staffordshire. He grew up in the village of Tettenhall, Wolverhampton and had two siblings, Tina-Louise Richmond (née Speight), and Jason Speight. His father, Oliver Warwick Speight, is a property developer, and his mother, Jacqueline Fordham Speight (née Parker), was an art teacher. Jacqueline died on 5 September 2008, aged 62. Speight attended the independent school Tettenhall College for a year, before moving to state comprehensive Regis School, now known as King's C.E. School, also in Tettenhall, at the age of 12. Speight stated in an interview he was a slow learner at school, with a short attention span, and art was a way for him to communicate. He said he did "very badly" because he was a victim of bullying, and the "daily ordeal for two years" forced him to become the "class joker". Speight left aged 16 and went on to attend Bilston Art School, where he took a degree in commercial and graphic art.

On the afternoon of 3 January 2008, Speight called emergency services after waking up to discover Natasha Collins's body in the bath at their St John's Wood flat, in north-west London. Speight told police that he and Collins had spent the previous evening "partying", drinking wine and vodka, and taking cocaine and sleeping pills. Speight was questioned by police and was subsequently arrested on suspicion of murder and of supplying Class A drugs, but was released on bail until the first week of February. Because of this, the BBC cancelled the Saturday repeat edition of SMart. An inquest, which opened on 8 January 2008, heard that the death was not thought to be suspicious but should be "subject to further investigation". At that point, police were awaiting results of toxicology tests after a postmortem examination was inconclusive. The BBC cancelled repeat broadcasts of SMart and SMarteenies until further notice, and on 28 February Speight announced he was quitting the show, because the "tragic loss" of Collins had left him unable to continue. Speight denied any involvement with Collins's death, and on 19 March it was reported that the police were no longer considering him a suspect.

In April 2008, the coroner recorded a verdict of death by misadventure in relation to Collins. The cause of death was "cocaine toxicity and immersion in hot water", according to the consultant pathologist. The inquest found that she had taken "very significant" amounts of cocaine with sleeping pills and vodka, and that she had suffered 60% burns to her body, including her tongue. The coroner noted that at some stage in the night after both Speight and Collins had gone to bed, Collins got up to have a bath. He said that it was "more likely than not" that a heart problem had caused Collins to fall unconscious while the hot tap was running. Following Collins's death, Speight moved in with Collins's mother.

On 13 April 2008, Speight's body was discovered hanging from the roof of MacMillan House, adjacent to London's Paddington Station, hidden from public view, six days after he died. The discovery was made by railway workers at 10:00 am, and British Transport Police confirmed that the body was Speight's on 14 April 2008. An inquest into his death opened on 16 April 2008, and a post-mortem confirmed the cause of death as hanging. It was then adjourned until 20 May. The police said Speight may have used a sixth floor fire exit to get to the area where he was found.

Speight's funeral was held on 28 April at St Michael and All Angels Church in Tettenhall and hundreds went to pay their respects. The service included a performance by the choir from Tettenhall College, Speight's former school, and his coffin was carried out of the church accompanied by the theme tune of SMart. He was later cremated and his ashes were interred in Tettenhall. In May, the inquest resumed and determined that Speight was deeply depressed by his fiancee's death. It was also disclosed that suicide notes had been found, one in his left pocket, and one addressed to his parents in his diary at his home. The notes described how he could not "contemplate life without [Collins]". The coroner, Dr Paul Knapman, said there were no suspicious circumstances surrounding the death. In May 2008, Speight's father created a foundation, Speight of the Art, or SP8 of the Art and launched it at a memorial service that took place on what would have been his 43rd birthday, 6 August 2008, at St Paul's Church in Covent Garden, London.

Speight was involved in charity work. He was President of the Muscular Dystrophy Campaign's Young Pavement Artists Competition, originally a one-off, year-long project that ended up lasting eight years, and he was a spokesperson for ChildLine. In 2007 he was the presenter of the Müller Big Art Project for Comic Relief in Trafalgar Square.

In 2004, Speight participated in Rolf Harris's Rolf on Art, for which a giant reproduction of John Constable's The Hay Wain was created in Trafalgar Square. In 2005, he was involved in a similar project where Hans Holbein's portrait of Henry VIII and Leonardo da Vinci's Mona Lisa were both reconstructed, the latter in the grounds of Edinburgh Castle. Speight had planned a project involving a trip to Borneo in March 2008 to train abused orangutans not to fight each other, but this never took place. Speight regularly toured with Speight of the Art, a series of art workshops he ran for children, and during the Christmas period, he performed in pantomime as "Buttons" in Cinderella at the Watersmeet, Rickmansworth, in December 2007.

Speight's other work ranged from children's television to adult factual programmes. His children's television credits include playing the Abominable No Man in Timmy Mallett's Timmy Towers and presenting Beat the Cyborgs, Name That Toon, On Your Marks, Insides Out, and History Busters, the last of which won a Royal Television Society Award. Speight also worked on This Morning, The Heaven and Earth Show, The Big Breakfast and was a contestant on ITV's Gladiators and Celebrity Wrestling. Speight also played the king on children's programme See It Saw It, where he met Natasha Collins. Collins was seriously injured after being hit by a car in 2001, and had to leave See It Saw It. Speight began dating her in 2003, and they became engaged in Barbados in 2005. They planned to get married in fancy dress and Speight joked that the wedding might feature monkeys, his favourite animal.

Speight intended to become a cartoonist, but he eventually became a TV presenter following a job painting the set of a television production. He auditioned for SMart and, following a successful interview where he met future co-presenter Jay Burridge, he went on to present SMart from its first edition in 1994. Speight became close friends with Burridge, whose art studio in West London was used to create all of the art content for SMart; Burridge noted: "We would bounce ideas and jokes off each other all day until we had developed an almost telepathically linked knowledge of what made each other laugh." Speight and Burridge were joined by third presenter Zoe Ball, who was replaced first by Josie d'Arby, and then Kirsten O'Brien and Lizi Botham. With Burridge, O'Brien and Botham, Speight presented the spin-off shows SMart on the Road, SMarteenies, and various live events. He starred in the BAFTA-nominated ITV Saturday morning show Scratchy & Co. from 1995 until 1998.

Mark Warwick Fordham Speight (6 August 1965 – 7 April 2008) was an English television presenter and host of children's art programme SMart. Speight was born in Seisdon, Staffordshire and left school at 16 to become a cartoonist. He took a degree in commercial and graphic art and, while working in television set construction, heard of auditions for a new children's art programme. Speight was successful in his audition and became one of the first presenters of SMart, working on it for 14 years.