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In October 2019, Dano was cast as The Riddler in Matt Reeves’s forthcoming 2021 film The Batman.

Dano made his directorial debut with the 2018 drama film Wildlife, based on the novel by Richard Ford and starring Carey Mulligan and Jake Gyllenhaal. He co-wrote the screenplay with his partner Zoe Kazan. In 2018 he starred in the Showtime miniseries Escape at Dannemora.

From December 27, 2018 to March 17, 2019, Dano starred in the critically acclaimed Broadway Revival of Sam Shepard's True West with Ethan Hawke at the Roundabout Theater Company’s American Airlines Theater in New York.

In the Autumn of 2016, he appeared in video as an onstage "stand-in" during the Nostalgia For the Present concert tour of Australian singer Sia Furler for her song, "Bird Set Free." In 2018, he portrayed escaped inmate David Sweat in the Showtime miniseries Escape at Dannemora alongside Patricia Arquette and Benicio Del Toro, for which he received a nomination for the Primetime Emmy Award for Outstanding Supporting Actor in a Limited Series.

In July 2016, it was announced that Dano would make his directorial debut with the movie Wildlife, based on the 1990 novel of the same name, by Richard Ford. The movie would be produced by June Pictures, and would star Carey Mulligan and Jake Gyllenhaal. Dano stated, "I have always wanted to make films and have always known I would make films about family. I couldn't be happier to have such beautiful collaborators like Carey and Jake leading the way." The film opened to rapturous reviews after its premiere at the Sundance Film Festival. It also played at the Cannes Film Festival, the Toronto Film Festival and the Mill Valley Film Festival. The film has earned a 94% on Rotten Tomatoes with the consensus reading, "Wildlife's portrait of a family in crisis is beautifully composed by director Paul Dano -- and brought brilliantly to life by a career-best performance from Carey Mulligan". Wildlife was nominated for 3 Independent Spirit Awards for Best First Feature, Best Female Lead (Mulligan), and Cinematography (Diego Garcia).

In 2015, Dano played a younger version of the Beach Boys founder Brian Wilson, with John Cusack as an older version of Wilson, in the biopic Love & Mercy, for which he received a Golden Globe Award nomination for Best Supporting Actor.

In 2015 Dano appeared with Michael Caine and Harvey Keitel in the Italian comedy-drama Youth; Dano portrayed Jimmy Tree, an actor who is researching for an upcoming role but is frustrated that he is best-remembered by the public for a prior role as a robot. In January 2016, Dano appeared as Pierre Bezukhov in the BBC's six-part adaptation of Tolstoy's War and Peace.

Dano has also received accolades for roles such as John Tibeats in 12 Years a Slave (2013) and Alex Jones in Prisoners (2013). His acting portrayal of musician Brian Wilson in Love & Mercy (2014) earned him a Golden Globe nomination in the category of Best Supporting Actor.

Dano appeared in three feature films in 2012: Ruby Sparks, as a writer whose fictional character (played by Zoe Kazan, the film's writer and Dano's girlfriend) inexplicably appears as a real person; time-travel thriller Looper, in a supporting role with Joseph Gordon-Levitt and Bruce Willis; and with Robert De Niro in Being Flynn as the film's writer Nick Flynn, about his relationship with his father. In 2013, Dano appeared in Steve McQueen’s period drama biopic 12 Years a Slave, based on the memoirs of Solomon Northup. Dano portrayed John Tibeats, an overseer at the plantation Northup is sold to. The film was a massive critical success and won the Academy Award for Best Picture, among numerous other awards.

In 2008, he starred in Gigantic, a poorly-reviewed film about a man seeking to adopt a Chinese baby, co-starring Zooey Deschanel. He reunited with Brian Cox in 2009's Good Heart, a low-budget English-language Icelandic film. He provided the voice of one of the creatures in the film adaptation of Where the Wild Things Are. He played a genius inventor in 2010's Knight and Day, an action thriller starring Tom Cruise and Cameron Diaz. The same year he appeared in Meek's Cutoff, a well-reviewed historical drama. In 2011, he had a supporting role in the big-budget science fiction film Cowboys and Aliens.

Dano appeared in several additional Broadway productions including A Thousand Clowns at the Roundabout Theatre, and in the Ethan Hawke directorial debut Things We Want during its 2007 Off-Broadway run.

Dano has been in a relationship with actress and screenwriter Zoe Kazan since 2007. They have a daughter, born in August 2018. They reside in Boerum Hill, Brooklyn.

He came to greater attention in 2006, when he played as Dwayne, a voluntarily mute teenager as part of an ensemble in the comedic drama Little Miss Sunshine, which received critical acclaim and collective awards for its cast. Dano had a dual role for the 2007 period film There Will Be Blood. This earned him a BAFTA nomination for Best Supporting Actor, and brought him positive reviews, with Texas Monthly saying that his performance was "so electric that the movie sags whenever he's not around." and Peter Travers remarking "All praise to the baby-faced Dano...for bringing sly cunning and unexpected ferocity to Plainview's most formidable opponent." Rolling Stone magazine included Dano in its Hot List for 2007, calling his performance style "Daniel Day-Lewis + Billy Crudup × Johnny Depp."

While he was a child, Dano's family moved to New Canaan, Connecticut, finally settling in Wilton, Connecticut. Dano continued his education there at Wilton High School, graduating in 2002 and attending Eugene Lang College in New York City. He was involved in community theater, and while he was performing in New Canaan, his parents were encouraged to take him to New York.

Dano acted in his first major film role when he was sixteen, playing Howie Blitzer, a teenage boy who becomes involved with a middle-aged ephebophile (Brian Cox) in L.I.E. (2001). He then appeared in the television film Too Young to Be a Dad as a high school student whose life is disrupted when his girlfriend becomes pregnant. In 2004, he played a small role as young Martin Asher in Taking Lives with Angelina Jolie and Ethan Hawke. In 2005, he played supporting roles in The King and The Ballad of Jack and Rose.

Dano started his career on Broadway before making his film debut in The Newcomers (2000). He won the Independent Spirit Award for Best Debut Performance for his role in L.I.E. (2002) and received accolades for his role as Dwayne Hoover in Little Miss Sunshine (2006). For his dual roles as Paul and Eli Sunday in There Will Be Blood (2007), he was nominated for the BAFTA Award for Best Supporting Actor.

At age ten, Dano was noted for roles in classical plays on Broadway, making his debut at age twelve in John Tillinger's Broadway revival of Inherit the Wind alongside George C. Scott and Charles Durning. He appeared in an episode of the sitcom Smart Guy and a minor role in the 2000 family drama The Newcomers, as well playing the part of Patrick Whalen, in a few episodes of The Sopranos in season 4.

Paul Franklin Dano (/ˈ d eɪ n oʊ / ; born June 19, 1984) is an American actor, director, screenwriter, producer, and musician.