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In April 2020, during the COVID-19 pandemic, Borland began publicly releasing previously unheard material from Eat the Day and Goatslayer, an experimental project he had with brother Scott since the two were teenagers. Borland claimed that the latter project had 23 albums worth of material, the first of which, "The Feather Serpent", was released on April 25.

Although originally from Richmond, Virginia, Borland spent his formative years growing up in Nashville, Tennessee. His initial interest in music took the form of drumming, but this ended quickly due to his parents disapproval of percussive instruments in the home. Borland then switched to taking guitar lessons from a member of his parent’s Presbyterian church. However, Borland quickly found his own musical tastes and interests at odds with local music trends which, due to his proximity to Nashville, skewed heavily towards country music. He later recalled, “I would bring in something, and my teacher would go, ‘I’ve never heard of The Damned. Don’t you want to play some Merle Haggard?’”

When Borland’s brother Scott took an interest in bass guitar, the two brothers began to play together. Borland found his interest in hip hop music piqued with the release of “Bring the Noise,” a collaboration between heavy metal band Anthrax and hip hop group Public Enemy.

Borland then relocated to Jacksonville, Florida with his parents where, disillusioned with the local music scene, he began attending the Douglas Anderson School of the Arts. While there, he explored other artistic endeavors, such as sculpture and special effects. However, Borland continued his pursuit of music through guitar lessons, working with a guitar teacher that specialized in jazz music. According to Borland, “My first teacher had ingrained playing by ear so much that, when my jazz teacher gave me a sheet of music and an audio tape to go with it, I would learn via the tape instantly, but I couldn’t make myself learn the sheet music.” As his guitar skills continued to improve, Borland began to craft his own guitar parts and style. Eventually, finding himself forced into church attendance and feeling a lack of interest in religion, Borland’s frustration and confinement pushed him to move out of his parent’s house at the age of 18.

In June 2017 he posted on social media that Big Dumb Face will be returning that year, eventually confirming that the new album, Where Is Duke Lion? He's Dead... would be released on October 31 via Edison Sound, with a full scale physical release to follow on November 17. Explaining why he revived Big Dumb Face after 16 years, he said that he wanted to make something "really heavy" in contrast to the post-rock of Crystal Machete, and that he was currently "into making things that [he] said [he'd] never do again", such as the solo record, or reviving Black Light Burns to release a new album in June 2018. He did, however, say that he would not tour with Big Dumb Face as he has prior touring commitments with Limp Bizkit and his ex-wife's band, Queen Kwong. In February 2018 Borland teased images of him working in the studio with Travis Barker on an "as yet untitled project". He also played guitar on five tracks from Jonathan Davis's debut solo album Black Labyrinth, which was recorded between 2007 and 2018.

On May 3, 2016, Borland released an instrumental solo album named Crystal Machete on his personal record label (Edison Sound), which saw him stepping outside his comfort zone by featuring no distorted guitars or vocals. The album was released to positive reviews, with Drowned in Sound calling the album a "a beautiful, sprawling post-rock mini epic". In 2016 he disbanded Black Light Burns.

In winter 2016 Limp Bizkit embarked on a co-headlining UK tour with Korn, including dates at Wembley Arena with support from Madball. As of October 2017, the status of the new Limp Bizkit album Stampede of the Disco Elephants is unknown, with Borland telling NME that he has completed all of his parts for "28 or 29 songs" and is waiting for Fred to choose what songs he wants to complete before he can mix and master it. He also blamed his time in Limp Bizkit for difficulty in getting jobs with other acts such as Nine Inch Nails and Marilyn Manson in the past, stating that the other bands didn't want to be associated with his previous band.

Borland is also known for performing in costumes and body paint during concerts, appearing in bunny and kung fu suits, and painted as a skeleton and what he describes as a "burnt match". Describing the character, he stated, "I go onstage wearing almost nothing. I have underwear and my boots on, and I paint my whole head black—from the neck up—and I have the black contacts. All you can see is these glowing teeth." Borland's black contacts were customized for him by a company noted for making contacts for the science fiction TV series Babylon 5.

Borland collaborated with Riot Games to develop Hybrid Worlds, the theme song for the League of Legends Season 3 World Championships. Borland performed live for the event on October 4, 2013, along with The Crystal Method, Tina Guo, and Black Light Burns alumni Danny Lohner and Joe Letz.

In 2012, Black Light Burns began performing after their brief hiatus and released their second studio album The Moment You Realize You're Going to Fall and a concept "album", Lotus Island, in January 2013. In the same year, during the Black Light Burns tour, Borland stated that he would never do a solo project. "If I would do a solo project I would put my name on the record. But that will never happen."

During the recording for Chocolate Starfish and the Hot Dog Flavored Water, Borland switched to 6-string guitars. The entire album was recorded using a Master Guitars Cremona semi-hollow body guitar, which is visible in the video for My Generation. However, Borland did not tour with this guitar and instead used Paul Reed Smith guitars, one of which is a Standard 24 in a Platinum Metallic finish, a Custom 24 in a Black Slate finish, and a 4-string baritone. After the tour, they were not seen again. A Facebook photo posted by Fred Durst in 2012 shows Borland playing the Cremona again for studio purposes.

In 2009, the original lineup of Limp Bizkit reunited and began touring, which resulted in a second Black Light Burns record being put on the backburner. Borland left Manson's employ in May of the same year, claiming his reasons for leaving included rejoining his old band and having creative differences with Manson, citing "It's the Marilyn Manson show over there." Limp Bizkit recorded a new album, which Borland named Gold Cobra. Released on June 28, 2011. It received mixed reviews, with multiple reviewers praising Borland's guitar playing. It peaked at No. 16 on the Billboard 200.

Borland also designed the album artwork for Fear and the Nervous System's 2011 eponymous debut album.

On August 2008, at the ETP Conference, it was announced that Borland joined Marilyn Manson as a guitarist. In the winter of 2008, Borland helped in the recording of The Color of Violence's 2009 album Youthanize as a bassist.

Black Light Burns released its début album, Cruel Melody in the spring of 2007 to critical acclaim. It featured Borland on the vocals and guitars. Allmusic writer Greg Prato wrote, "For many, Borland was the only Limp Bizkit member who was taken seriously. As evidenced by Cruel Melody, he is now officially allowed to follow whichever musical path he so desires.". In 2008, Borland, along with Richard Fortus and Sugizo, supported Japanese metal band X Japan at their sold out reunion concerts at the Tokyo Dome.

In 2005, Yamaha approached Borland about a new signature guitar, which was almost 100% designed by Borland. The model CV820WB was released that same year. A semi-hollow body guitar with a large body and new Yamaha high-output split field humbuckers, made exclusively for that guitar, it also came with the Yamaha Quick Change finger-clamp locking tremolo system, which rids the user of having to cut the ball ends off of the strings, which is very unusual for a locking tremolo. This was Borland's main guitar for the recording of Black Light Burns' Cruel Melody and the touring behind it, as well as most of the touring Limp Bizkit did before the recording of Gold Cobra. Despite its innovative characteristics, it was not a popular guitar, and was discontinued in mid-2011, when Borland switched to Jackson Guitars.

After the 'failure' of another project, The Damning Well, in 2003, Borland began writing a solo record with its members Danny Lohner and Josh Freese, with Josh Eustis and Jonathan Bates contributing. This record served as the precursor to Black Light Burns. In August 2004, Borland rejoined Limp Bizkit and recorded another album, The Unquestionable Truth (Part 1) in 2005. Following the release of the band's Greatest Hitz album, the band went on hiatus after arguments broke out on MySpace between Borland and frontman Fred Durst, with Borland stating, "As of right now, none of my future plans include Limp Bizkit." In 2005, Borland formed Black Light Burns. Borland stated that this was now his main project, and anything else he did, including Limp Bizkit, was a side project. Borland toured with From First To Last roughly since early 2006. He announced planned for a fall 2006 tour that never went through due to Black Light Burns needing to find a new record label after Borland left Geffen Records. Borland had discussed plans to write and perform on the next From First To Last album, but he left the band when Black Light Burns' busy schedule started to pick up, leaving no room to work with From First to Last.

In March 2001, Big Dumb Face released its début album, Duke Lion Fights the Terror!!. Borland stated that the band's music is "really silly and idiotic and bizarre. [...] It's nothing but stupid [...] just all these retarded songs." In the fall of 2001, Borland left Limp Bizkit citing creative differences with the band, and formed the band Eat the Day, which he eventually abandoned due to being unable to find a vocalist. The unfinished demos from Eat the Day were subsequently released in April 2020 by Borland.

Significant Other saw Limp Bizkit reaching a new level of commercial success. The album climbed to No. 1 on the Billboard 200, selling 643,874 copies in its first week of release. In 2000, Durst announced that the band's third studio album would be titled Chocolate Starfish and the Hot Dog Flavored Water. The press thought that Durst was joking about this title. The album title is intended to sound like a fictional band; the phrase "Chocolate Starfish" referred to Durst himself, as he had frequently been called an asshole. Borland contributed the other half of the album's title when the band was standing around at a truck stop, looking at bottles of flavored water, and Borland joked that the truck stop didn't have hot dog or meat-flavored water. It debuted at number one on the Billboard 200, selling 1.05 million copies in its first week, and was the 18th best-selling album of the 2000s in the US.

Borland uses Ernie Ball .011-.052 gauge nickel-wound strings, where the G string is wound. For Limp Bizkit's first three albums, Borland used Mesa-Boogie Dual and Triple Rectifier amps. In 2000, Borland would use the Mesa Boogie heads in conjunction with a Diezel VH4. In the mid 2000s, on the recording of Cruel Melody and The Unquestionable Truth (Part 1), Borland's main amp was the Diezel VH4. Starting with the 2008 recording of The Moment You Realize You're Going to Fall Borland played mainly Orange Amps (specifically the Thunderverb 100 models) since touring in support of Cruel Melody began and has continued to use them since then, also on Gold Cobra. Currently, Borland plays EVH Amps and cabinets for his heavier tones. The mainstay of Borland's amp setup has been a Roland JC-120 combo amplifier to generate his particularly exceptional clean tones.

In a 1999 Spin profile on Limp Bizkit, when asked "Where will you be ten years from now?", Borland responded, "Probably not in the band."

Although the album was met with minimal response, touring consistently increased Limp Bizkit's success, and the third single from Three Dollar Bill, Yall, "Faith", became a radio hit, leading to a slot on Ozzfest, a tour organized by Ozzy and Sharon Osbourne. In 1998, Borland formed a side project, Big Dumb Face with his brother, influenced by Ween and Mr. Bungle.

In April 1998, Wes married his long-time girlfriend Heather McMillan, but later divorced in 2001. His second marriage was with Anna Carlise in March 2009 but they divorced in 2013. In October 2016 he married Carré Callaway, lead singer of Queen Kwong. The couple resided together in Detroit. She filed for divorce in 2019.

Borland used Ibanez seven-string guitars exclusively from 1997 until 1999. However, he stated that he played a 6-string guitar on the entire Three Dollar Bill, Yall record. Shortly after embarking on a tour with Korn, Ibanez got a hold of Borland and gave him a number of seven-string Universe guitars, essentially for free, which he continued to use extensively. During the touring in support of Significant Other, Borland used two custom Ibanez RG seven-strings with the electronic setup of an Ibanez AX7521 (two volume knobs and two tone knobs rather than one volume and one tone). Nothing is certain about the type of pickups that he used, however, he did use EMG pickups at one point, stating in an interview with GuitarCenter that he was moving from "passive to active pickups". Borland also endorsed the rare Ibanez RG7 CST guitar, which is made from superior/high-quality materials and is also equipped with an L.R. Baggs designed piezo system on a locking tremolo. He is currently one of two American owners of one. Additionally, Borland used a vintage Ibanez Musician MC150PW, which was modified to be fitted with 4-strings, which he used on songs such as "Nookie", "Full Nelson", and "The One". This guitar can be seen in the music video for "Nookie." Ibanez then made him a custom baritone 4-string AX that would be used to replace the Musician. This was eventually replaced by a custom-made Master Guitars "Cremona" 4-string, which he uses to this day.

He gained popularity when Limp Bizkit achieved mainstream success in the late 1990s and early 2000s. He formed Big Dumb Face with his brother Scott in 1998 and left Limp Bizkit in 2001 to started side projects including Eat the Day and The Damning Well. After rejoining Limp Bizkit in 2004, Borland founded Black Light Burns, with whom he has released three studio albums and a covers album. Limp Bizkit went on hiatus following the release of their album The Unquestionable Truth (Part 1) (2005). However, the band's original lineup reunited in 2009 and recorded their fifth studio album, Gold Cobra (2011). In 2016, Borland released his solo album Crystal Machete.

Borland was endorsed by Jackson shortly before the release of Gold Cobra, and owns several Rhoads guitars, which he uses on Limp Bizkit shows. In addition he plays various Warriors and King Vs, one of which is a left-handed black King V with white bevels that was converted by Borland into a right-handed guitar. All of these instruments are equipped with a Floyd Rose and a single Seymour Duncan Invader pickup in the bridge position. He does not currently have a signature model with Jackson. Additionally, Borland also uses a Fender Bass VI, tuned to one octave below E standard tuning. He uses this on "The Story" on The Unquestionable Truth (Part 1), and again on "90.2.10", "Walking Away", and "Back Porch" on the Gold Cobra record.

Borland tunes his guitars to C# standard tuning (C# F# B E G# C#) and to Drop B tuning (B F# B E G# C#). He also tunes his 4-string baritone guitars to a variant of this tuning with a low F# string, which is a bass string (F# F# B E). During the late 1990s when Borland played 7-string guitars, he played them like one would play a 6-string guitar by tuning the highest string to C# as well, while maintaining standard C# tuning all the way to the seventh string. This technique was later used by Stephen Carpenter of Deftones so he could play the songs he originally recorded on a six-string without losing the feel of a seven-string, as Limp Bizkit and the Deftones had toured together in the mid 1990s.

Other guitars Borland has been seen with before are a 1976 Fender Starcaster a stock ESP LTD V-401DX (used on the Limp Bizkit reunion tour for playing songs on Results May Vary) and Mayones Regius Pro 6, Legend and Setius GTM 6 Baritone guitars.

Wesley Louden Borland (born February 7, 1975) is an American rock musician and artist. He is best known as the current guitarist and backing vocalist of the American rap rock band Limp Bizkit, the lead vocalist and guitarist of the alternative and industrial rock band Black Light Burns, and the co-founder of metal band Big Dumb Face.